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Warp Darkmatter attacks a strange planet with Crater of Banpanaliya that holds a secret: it contains a secret gas that devolves everyone who smells it. As a result, the protectors become devolved into lemur-like creatures, Buzz gets devolved into a caveman, Mira becomes blue sludge and Booster becomes a dinosaur-like creature, leaving only XR to fight Warp and Zurg's Hornets.
- Patrick Warburton as Buzz Lightyear, LGMs
- Stephen Furst as Booster
- Nicole Sullivan as Mira Nova
- Diedrich Bader as Warp Darkmatter
- Jeff Bennet as Binipinardian #2 & #3
- Jim Cummings as Binipinardian #1, Announcer
- Neil Flynn as XR
- Wayne Knight as Zurg
- Frank Welker as Binipinardians